Well, I just tried those export settings, and it gives me the same error. But I turns out I forgot to press the export animation tab, so I was exporting without animations. It appears exporting with animations gives the error, no matter the settings.
After some looking, I think I've figured out the problem. One of your animations (Start) does not have save this data block even if it has no users (the F button beside the number of instances) and that seems to be causing the issue. It appears all of your animations have to be force saved in order to export.
Here's a picture showing what the animation(s) should look like (with the F button highlighted):
Once all of the animations had save this data block even if it has no users, I was able to export with the settings above.
Side note: Whenever I've exported with Apply Modifiers I've found it does not export my animations in the .dae file due to the Armature modifier being applied. If your exported file does not have animations, you'll probably want to try exporting with Apply Modifiers disabled/unchecked.
